Gray-cheeked Thrushes forage by consuming fruit in trees (or on the ground) and taking invertebrate prey on the ground with a quick peck.
They breed in willow-alder thickets and dense undergrowth in stands of spruce, balsam fir, other conifers, or cottonwood trees.
Migrants and wintering birds occupy many habitat types, but tend toward well-developed understories and berry bushes.
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